Haplogroup R1a-M420 originated ca. 25000 BC, and has been proposed to diverge initially in the vicinity of present-day Iran based on a study of modern populations[Underhill et al. 2015].
Hunter-gatherers of R1a-M420 lineage were proposed to have migrated from the Iranian area to the forests of Eastern Europe, since early samples were found in the Narva culture[Horvath 2015].
- [Horvath 2015] ^ Horvath, Csaba Barnabas. 2015. R1a sublcades and Bronze Age migrations on the Eurasian steppes. ESJ 2.
- [Underhill et al. 2015] ^ Underhill, P. A., G. D. Poznik, S. Rootsi, M. Jarve, A. A. Lin, J. Wang, B. Passarelli, J. Kanbar, N. M. Myres, R. J. King, J. Di Cristofaro, H. Sahakyan, D. M. Behar, A. Kushniarevich, J. Sarac, T. Saric, P. Rudan, A. K. Pathak, G. Chaubey, V. Grugni, O. Semino, L. Yepiskoposyan, A. Bahmanimehr, S. Farjadian, O. Balanovsky, E. K. Khusnutdinova, R. J. Herrera, J. Chiaroni, C. D. Bustamante, S. R. Quake, T. Kivisild, and R. Villems. 2015. The phylogenetic and geographic structure of Y-chromosome haplogroup R1a. Eur J Hum Genet 23 (1):124-31.