A more recent, revised and updated version of this paper has been published (2019)
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R1a-M420 and East Hunter-Gatherers

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Haplogroup R1a-M420 originated ca. 25000 BC, and has been proposed to diverge initially in the vicinity of present-day Iran based on a study of modern populations (Underhill et al. 2015). Hunter-gatherers of R1a-M420 lineage were proposed to have migrated from the Iranian area to the forests of Eastern Europe, since early samples were found in the Narva culture (Horvath 2015).

References

Horvath, Csaba Barnabas. 2015. R1a sublcades and Bronze Age migrations on the Eurasian steppes. ESJ 2.

Underhill, P. A., G. D. Poznik, S. Rootsi, M. Jarve, A. A. Lin, J. Wang, B. Passarelli, J. Kanbar, N. M. Myres, R. J. King, J. Di Cristofaro, H. Sahakyan, D. M. Behar, A. Kushniarevich, J. Sarac, T. Saric, P. Rudan, A. K. Pathak, G. Chaubey, V. Grugni, O. Semino, L. Yepiskoposyan, A. Bahmanimehr, S. Farjadian, O. Balanovsky, E. K. Khusnutdinova, R. J. Herrera, J. Chiaroni, C. D. Bustamante, S. R. Quake, T. Kivisild, and R. Villems. 2015. The phylogenetic and geographic structure of Y-chromosome haplogroup R1a. Eur J Hum Genet 23 (1):124-31.